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A more direct method of finding out about the variety of palaeofaunal disease is to turn to the palaeopathology of the animals concerned. During the last two hundred years the palaeopathology of Quaternary extinct fauna has been studied at some length elsewhere (Esper 1774; Goldfuss 1810; Schmerling 1835; Mayor 1854; Moodie 1917, 1923; Pales 1930). In fact, it is widely recognised that the formative period of palaeopathological investigation, from 1774 until the 1870s, largely concentrated on the disease of animals (Pales 1930; Roney 1966; Armelagos et al 1971; Chaplin 1971; Siegel 1976; Wells and Lawrance 1976; Ubelaker 1982).

Like most of the conditions described here psittacosis was most likely confined to isolated outbreaks and, in terms of the unique incidence among koalas, it is interesting to speculate if any of the extinct megafauna species carried this disease. Nevertheless, it is almost certain that another chlamydial species ( C trachomatis) existed in Australia at least 12,000 years ago. The evidence for this has been described elsewhere but manifests itself as scarring in the orbits of fossil human crania from southeastern Australia dated to that time (Webb 1990a).

And Staphylococcus sp. (Mims 1977). Salmonella paratyphosa has been isolated in primates, which may tell us something about the zoonotic origins of paratyphoid and the possibility for the ancient hunters of these animals to contract it. Others have pointed out that a number of organisms which live in the intestinal tract, including those causing dysentery (Shigella dysentariae), may well have existed in early huntergatherer societies (Hare 1967). We may remember that both Shigella and Salmonella are organisms known to cause infectious or septic arthritis among these societies (Ortner and Putschar 1981).

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